In order to study the galloping mechanism of ice-covered four-split transmission lines in oblique flows, the aerodynamic forces and aero-elastic response characteristics of the crescent-shaped four-split ice-covered transmission lines are investigated through wind tunnel tests on rigid and aero-elastic models. According to Den Hartog and Nigel’s galloping theories, the damping coefficients are calculated based on the experimental data. The results show that the crescent-shaped ice-covered four-split transmission lines usually suffer from torsional galloping. Furthermore, based on the aero-elastic wind tunnel data, the galloping is characterized by an elliptical trajectory, negative damping ratio, and a negative strain at hanging position. In addition, the galloping appears to be more prone to occur under oblique flows, with a larger galloping amplitude and a lower critical wind speed. This might be because an out-of-plane vibration of the third-order mode is excited at a lower wind speed, leading to a coupled resonance between in-plane and out-of-plane vibrations at the third-order mode with a frequency ratio of 1:1. The experimental results in this paper can also be used to verify the fluid-structure interaction simulation method of ice-covered transmission lines.