“…Invertebrate movements involved in colonisation and redistribution are normally attributed to one of four possible mechanisms; downstream drift, aerial sources, upstream and vertically directed movements (Williams & Hynes, 1977), but random lateral movements can contribute a fifth mechanism, especially when new areas, such as implanted substrates, become available within populated habitats. Downstream drift is usually considered to be the most important colonisation/redistribution mechanism (Townsend & Hildrew, 1976;Bird & Hynes, 1981;Graesser & Lake, 1984), although the propensity to drift is not the same in all species (Chaston, 1972;Turcotte & Harper, 1982;Kohler, 1983;Allan, 1984). Other studies have demonstrated that movement on, or in, the substrate can also contribute significantly to overall redistribution of benthos (e.g.…”