Differentiation and allozyme variability of shads genus alosa (Clupeiformes, Alosiinae) from Azov-Black Sea basin

Delimitation of Alosa species (Teleostei: Clupeiformes) from the Sea of Azov: integrating morphological and molecular approaches

“…The uncertain taxonomic status of Alosa spp. from the Black Sea and Sea of Azov has been previously reported in a few studies that examined morphological and allozyme diversity within the Alosa species complex and it was suggested that the observed variation can be explained by intraspecific plasticity and that multiple species described from the Black Sea and Sea of Azov basin might better be considered morphotypes of a single species (Mezhzherin et al, ; Pravotorov, ).…”

“…The first two species are endemic to the Black Sea and Sea of Azov, while the third was originally described from the Caspian Sea. A subspecies of A. caspia, Alosa caspia tanaica (Grimm, 1901), has been described from the Sea of Azov, but the taxonomic status of A. c. tanaica is still debated (Hoestlandt, 1991;Kottelat, 1997;Kottelat & Freyhof, 2007;Mezhzherin et al, 2009;Svetovidov, 1952Svetovidov, , 1964Whitehead, 1985). In the original descriptions, distinctions between species were based on the number of fin rays, colouration and body proportions (i.e., eye diameter, body height, head length and head height).…”

Marine fishes in the Black Sea: recent conservation status

“…A(M) DD 3, 5 Abaza et al, 2006;Fricke et al, 2007; Spicara smaris (Linnaeus, 1758) A(M) NE 1, 2, 3, 5 Fricke et al, 2007;Keskin, 2010;Karapetkova & Zhivkov, 2010;Zivkov et al, 2005;Spicara maena (Linnaeus, 1758) A(M) NE 1, 2, 4, 5, 6 Boltachev & Karpova, 2012;Fricke et al, 2007;Keskin, 2010;Karapetkova & Zhivkov, 2010;Zivkov et al, 2005; Spicara flexuosa Rafinesque, 1810 2, 3, 4, 5, 6 Boltachev & Karpova, 2012;Karapetkova & Zhivkov, 2010;Salekhova, 1979;Vasill'eva, 2007;Zivkov et al, 2005; Clupeidae Alosa immaculata Bennett, 1835 2, 3, 4, 5, 6 Dobrovolov et al, 2012, Fricke et al, 2007Freyhof & Kottelat 2008;Keskin, 2010;Karapetkova & Zhivkov, 2010;Mezhzherin et al, 2009;Turan et al, 2010;Vasil'eva, 2007; Alosa caspia (Eichwald, 1838) E(B) LC 1, 2, 3, 4, 6 Dobrovolov et al, 2012;Karapetkova & Zhivkov, 2010;Mezhzherin et al, 2009;Vasil'eva, 2007;Zivkov et al, 2005 2, 3, 4, 5, 6 Chashchin, 1998;Dobrovolov, 1976;Fricke et al, 2007;Grant, 2005;Ivanova & Dobrovolov, 2006;Ivanova et al, 2013a;Keskin, 2010;Karapetkova & Zhivkov, 2010;…”