“…The partitioning of water above and below the surface is important for a wide range of Earth-surface processes: the extent and connectivity of surface flows determine the mobility and dispersion of freshwater macroinvertebrates (Clarke et al, 2008); fast-moving surface water is required to move sediment and carve valleys into the landscape (Howard et al, 1994); and the total land area covered by surface water is a key control on carbon-dioxide efflux from continents, as well as dissolved organic carbon (DOC) export in rivers Aufdenkampe et al, 2011;Zimmer & McGlynn, 2018). However, even within an individual catchment, the density of actively flowing streams can vary by up to an order of magnitude seasonally or during rainstorms Blyth & Rodda, 1973;Day, 1978;Godsey & Kirchner, 2014;Goulsbra et al, 2014;Gregory & Walling, 1968;Jensen et al, 2017;Lovill et al, 2018;Roberts & Archibold, 1978;Roberts & Klingeman, 1972;Shaw, 2016;Whiting & Godsey, 2016;Wigington et al, 2005;Zimmer & McGlynn, 2017; Figure 1), and it is unclear what controls the temporal variability in wetted stream length. However, even within an individual catchment, the density of actively flowing streams can vary by up to an order of magnitude seasonally or during rainstorms Blyth & Rodda, 1973;Day, 1978;Godsey & Kirchner, 2014;Goulsbra et al, 2014;Gregory & Walling, 1968;Jensen et al, 2017;Lovill et al, 2018;Roberts & Archibold, 1978;Roberts & Klingeman, 1972;Shaw, 2016;Whiting & Godsey, 2016;Wigington et al, 2005;Zimmer & McGlynn, 2017; Fi...…”