“…Among larger karst streams, regionally endemic fishes are generally concentrated in the stenothermal, headwater sections of streams, whereas fishes with more ubiquitous distributions are generally concentrated in the more abiotically variable, downstream sections of streams (Hubbs, 1995). Because of their limited distributions in or near a spring source, these regionally endemic fishes are often described as spring-associated; however, exact factors leading to their restricted range as well as their segregation with riverine taxa remain unclear (Garrett et al, 2004). Several biotic (e.g., competition, fecundity) and abiotic (e.g., temperature, salinity, stream order, stream gradient) factors are hypothesized to regulate fish distributions and segregation among fish assemblages (Garrett et al, 2004;Gido et al, 1999;Ingersoll and Claussen, 1984;Taylor and Lienesch, 1996).…”