“…The significance and importance of coastal dune systems are well known (Arens et al, 2001;Hanley et al, 2014;Ruz et al, 2009). Coastal sand dunes, and particularly foredunes, act as a protective barrier on many of the world's coastlines (Carter, 1991;Doody, 2012;Maximiliano-Cordova et al, 2019;Simm et al, 1996) and, hence, are an important natural defence against inundation by seawater of economically and environmentally significant land, infrastructure, and activities (Fernández-Montblanc et al, 2020;Hanley et al, 2014;Maximiliano-Cordova et al, 2019). They are also intrinsically areas of high ecological significance themselves, as well as sources of raw materials, grazing land, and recreation (Doody, 2012;Everard et al, 2010;Hanley et al, 2014;Martínez et al, 2007).…”