“…Covert reproduction by delayed dispersers during forays would provide evidence of a previously undocumented pathway to fi tness, and suggests that delaying and foraying may be more of an alternative life-history strategy than simply "making the best of a bad situation," as some 391 have suggested (e.g., Emlen 1997). Further, if covert extraterritorial helper reproduction is common in many species, the behavior even has the potential to alter perceptions about the costs and benefi ts of cooperative breeding and delayed dispersal, which have been debated for several decades (Skutch 1935, Brown et al 1978, Vehrencamp 1980, Brown 1987, Stacey and Koenig 1990, Gerlach and Bartmann 2002, Haydock and Koenig 2003, Griffi n et al 2003. A molecular genetic investigation of parentage in Micronesian Kingfi shers has the potential to lend insight into this phenomenon.…”