“…They are known as the primary carbon storage in foraminifera (Hottinger & Dreher, 1974, Leutenegger, 1977, Hottinger, 1982, Pawlowski et al, 1995. Similarly to residual bodies, the accumulation of lipid droplets in foraminifera is a known symptom of environmental stress, such as heavy metal toxicity and oil pollution (Morvan et al, 2004, Le Cadre & Debenay, 2006, Frontalini et al, 2015, Frontalini et al, 2018. Our results show that starvation can be another factor triggering lipid droplet accumulation.…”