The ontogeny of throwing and striking R i c h a R d W . Y o u n g r E V i E W abstract humans are the most adept throwers in the animal kingdom. although throwing has been recorded in other primates, it is rare, weak, inaccurate and entails only arm movement. in contrast, human throwing is an explosive act employed from a bipedal stance that involves a complex, exceedingly rapid, full-body movement. This motion generates a pulse of kinetic energy in the legs that is progressively augmented by the pelvis, trunk, arm and hand, where it is transferred to a missile that is launched toward the target with high velocity and accuracy. Linking the study of throwing development in young children with analysis of the adult throwing motion makes it possible to address the question of whether human throwing is an acquired behavior or is based primarily on an inherited motor program. current evidence supports the conclusion that the role of instruction and learning is minimal. human throwing is predominantly the result of an innate motor program which emerges at a very early age in all children without teaching, yields a throwing motion that is the forerunner of the one used by adult athletes, is characterized by a prominent gender difference, and proceeds in some adults to a high level of proficiency. The same conclusions apparently apply also to striking (club-swinging), which employs a similar full-body motion. an explanation for these observations is presented, based upon the concept that throwing and club-swinging prowess provided reproductive advantages during early human evolution.introduction Darwin (1871) was the first to call attention to the remarkable ability of our species to throw with speed and accuracy. "To throw a stone with as true an aim as can a Fuegian in defending himself, or in killing birds," he wrote, "requires the most consummate perfection in the correlated action of the muscles of the hand, arm, and shoulder, not to mention a fine sense of touch" (1871, p. 138). Such behavior is rarely recorded in other primates, and when observed it is ineffective. darwin associated human throwing prowess with a distinctive body structure. The hands and arms could hardly have become perfect enough to have hurled stones and spears with a true aim, he believed, as long as they were habitually used for locomotion, body support, or climbing trees; furthermore, the successful thrower must stand firmly on his feet in an upright, bipedal stance. Today, darwin's conclusions are thoroughly documented. Modern human adults are indeed the most effective throwers by a wide margin, a generalization that can be expanded to include the related behavior of swinging clubs. in their highest development, both motor skills employ a rapid, coordinated sequence of muscle contractions that begins in the legs and traverses the body, progressively accumulating kinetic energy that ultimately is transmitted to the hand, which then projects the missile or club towards a target.is this behavior inherited or learned? is it based upon an innat...