This paper focuses on sense of place as a cultural ecosystem service and a mediatory experience in grasping a collection of ecosystem benefits. Through phenomenological and social constructionism/relational approaches, we focus on sense of place as a relational entity at both the individual and collective levels. Using content analysis of 32 in-depth interviews with (a) environmental officials and (b) local inhabitants in the region of Ostrobothnia, Finland, the study finds that sense of place is shaped via the interactions of environmental settings and cultural practices. Here, we argue that material and non-material interactions contribute to sense of place. With a phenomenological approach, the study presents the qualities associated with places, such as tranquility, beauty, and uniqueness that are important for achieving both material and non-material ecosystem benefits, while social constructionism/relational approaches indicate that the social dimension of sense of place contributes to ecosystem service benefits through traditional cultural practices. The study also shows a shift in cultural practices from provisioning to recreation in creating meaningful places. The results highlight a multilocal dimension of senses of places, particularly in contemporary lifestyles and increased mobilities.