“…Tropical rainforest in Borneo is experiencing threats from rampant forest conversion, selective logging, hunting for bushmeat, forest fire and wildlife trade (Taylor et al, 1999;Bennett et al, 2002;Kinnaird et al, 2003;Sodhi et al, 2004;Nakagawa et al, 2006;Linkie et al, 2007;Bernard et al, 2009;Gaveau et al, 2014;Brodie et al, 2015a). The proliferation of road construc-tions and expansion in Sarawak in an unprecedented manner jeopardises forest connectivity, fragmenting forests into patches and catalyses anthropogenic activities within and around these areas (Clements et al, 2014;Alamgir et al, 2020). Forest gaps and fragmentation created by roads in forests would, in turn, induce a negative impact on population density, diversity, disruption to animal movements and their daily activities (Laurance et al, 2009;Tsuyuki et al, 2011;Clements et al, 2014;Mohd-Azlan et al, 2018a).…”