“…For example, while more productive environments house more species, tropical ecosystems house more diversity at all scales than would be expected by environmental production alone 15,26 . Similarly, studies of human cultures show that while diversity tends to be higher in more productive environments 1,3,4,6,23,24,40 there are many other environmental, climatic, geological, and cultural processes that impact diversity at various scales [40][41][42][43][44][45] . For example, while global variation in hunter-gatherer population density, space use, and mobility is well-predicted by environmental productivity 44,46,47 , the spatial distribution of ethnolinguistic groups in general across the socioeconomic spectrum is better predicted by their level of sociopolitical complexity 45,48 .…”