“…We found that a lowtemperature treatment significantly damaged the chloroplast structure, loosening the thylakoids until they were disintegrated and increasing the number of liposomes in the chloroplasts, indicating that the photosynthetic apparatus was destroyed in both species during the late stage of the lowtemperature stress. Similar observations have also been made (Taylor and Craig, 1971;Marzanna et al, 2002). Thus, we can conclude that the decrease in the rate of photosynthesis during the late stage of low-temperature stress was mostly due to non-stomatal factors that damaged the photosynthetic mechanism by analysing results above.…”