We present observations of three distinct transits of HD 17156b obtained with the Fine Guidance Sensors (FGS) on board the Hubble Space Telescope (HST ) 5 . We analyzed both the transit photometry and previously published radial velocities to find the planet-star radius ratio R p /R ⋆ = 0.07454 ± 0.00035, inclination i = 86.49 +0.24 −0.20 deg, and scaled semi-major axis a/R ⋆ = 23.19 +0.32 −0.27 . This last value translates directly to a mean stellar density determination ρ ⋆ = 0.522 +0.021 −0.018 g cm −3 . Analysis of asteroseismology observations by the companion paper of Gilliland et al. (2009) provides a consistent but significantly refined measurement 6 of ρ ⋆ = 0.5308 ± 0.0040 g cm −3 . We compare stellar isochrones to this density estimate and find M ⋆ = 1.275 ± 0.018 M ⊙ and a stellar age of 3.37 +0.20 −0.47 Gyr. Using this estimate of M ⋆ and incorporating the density constraint from asteroseismology, we model both the photometry and published radial velocities to estimate the planet radius R p = 1.0870 ± 0.0066 R J and the stellar radius R ⋆ = 1.5007 ± 0.0076 R ⊙ . The planet radius is larger than While this work was in press, the companion work revised this measurement to ρ ⋆ = 0.5301 ± 0.0044 g cm −3 . We note that the change, which is less than 0.2 of the error, would translate to a stellar radius change of less than 0.001 R ⊙ and does not materially affect the results presented in this paper.that found in previous studies and consistent with theoretical models of a solarcomposition gas giant of the same mass and equilibrium temperature. For the three transits, we determine the times of mid-transit to a precision of 6.2 s, 7.6 s, and 6.9 s, and the transit times for HD 17156 do not show any significant departures from a constant period. The joint analysis of transit photometry and asteroseismology presages similar studies that will be enabled by the NASA Kepler Mission.