“…15–13 Ma) with the establishment of a permanent Antarctic ice‐sheet (Zachos et al, 2008; Liu et al, 2009; Beddow et al, 2016; Hutchinson et al, 2021). These periods coincided with the emergence of seasonal biomes in Europe and Asia, the aridification of Australian, African, and Asian inlands, and the subsequent contraction of the tropics with tropical forests retreating to lower latitudes (Morley, 2003; Bowen, 2007; Contreras et al, 2013; Fang et al, 2015; Lin et al, 2015; Beasley et al, 2021; Couvreur et al, 2021). Moreover, all above geo‐climatic changes, associated with sea level drops, have led to major turnovers of the vegetation (Sun et al, 2015; Pound & Salzmann, 2017; Couvreur et al, 2021), with successful plant dispersal across different paleogeographic regions (Jiang et al, 2019), and the establishment of present‐day genera and families in multiple regions, probably pre‐adapted to current climates (e.g., Linnemann et al, 2017; Su et al, 2018; Huang et al, 2021; Ling et al, 2021).…”