The Chalk is an unusual karst aquifer with limited cave development, but extensive networks of smaller solutional conduits and fissures enabling rapid groundwater flow. Small-scale karst features (stream sinks, dolines, dissolution pipes, and springs) are common, with hundreds of stream sinks recorded. Tracer velocities from 27 connections between stream sinks and springs have median and mean velocities of 4700 and 4600 m.d.−1. Tests to abstraction boreholes also demonstrate very rapid velocities of 1000s m.d.−1. Natural gradient tests from observation boreholes have rapid velocities of 100s m.d.−1. There is strong geological control on karst with dissolution focused on stratigraphical inception horizons. Surface karst features are concentrated near the Palaeogene boundary, or where thin superficial cover occurs, but rapid groundwater flow is also common in other areas. The Chalk has higher storage and contaminant attenuation than classical karst, but recharge, storage and flow are influenced by karst. Point recharge through stream sinks, dolines, losing rivers, vertical solutional fissures, and soakaways enables rapid unsaturated zone flow. Saturated zone networks of solutional fissures and conduits create vulnerability to subsurface activities, and enable long distance transport of point source and diffuse pollutants, which may be derived from outside modelled catchment areas and source protection zones.