“…Tea plant leaves produce many polygalloylated glucose derivatives ( Yang and Tomás-Barberán, 2018 ; Wei et al, 2019 ). β-Glucogallin is generated from UDP-glucose and gallic acid in tea plants by UDP-glucose: galloyl-1-O-β-D-glucosyltransferase CsUGT84A22 ( Liu et al, 2012 ), whose homologs were also reported in several other plants ( Ono et al, 2016 ; Tahara et al, 2018 ; Zhao et al, 2020 ). β-Glucogallin is further used as the galloyl donor in the sequential galloylation of 1,6-di-O-galloyl-β-D-glucose to generate 1,2,6- tri-, 1,2,3,6- tetra-, and 1,2,3,4,6-penta-O-galloyl-β-D-glucose ( Niemetz and Gross, 2005 ).…”