This paper provides an introduction to the ecological fabric of northern Australia, described here as being a land characterised by extreme climatic seasonality and largely devoid of marked topographic features. Largely as a result of the latter trait, many species have extensive geographic ranges, and the spatial turnover in species composition is extremely limited. Somewhat counter-intuitively, these two traits can be accommodated by organisms only through reliance on critical, but often subtle, landscape variation. We present some preliminary models for Gouldian finch (Erythrura gouldiae) and black-footed tree-rat (Mesembriomys gouldii) to illustrate patterns of variation in their resource availability, and the consequences of such variation. We discuss briefly some studies that have attempted to integrate, or at least consider, these elements.