This article is focused on problematic distinctions of difference among animals in the lineage of great apes. It combines several theoretical perspectives on evolutionary relationships, technological innovation, the development of body parts as tools, and a semiotic interpretation of what André Leroi‐Gourhan called technicity. Foundational questions in social theory are developed using biosemiotics, particularly as concerns a materialist understanding of religion and the magical aspects of cultural representation. This, it is argued, provides a framework for theorizing social history in terms of real ecological relations, embodied meaning, and the transference of meaning onto objects. Understood semiotically, the material history of Hominidae, encompassing animals with different kinds of motility, dexterity, and techno‐semiotic orientations towards the world, is inclusive and relational rather than exclusively anthropocentric, as is the case for social theory based on the artifice of language and articulations of belief, creativity, and cultural distinction that are thought to be distinctive of the genus Homo.