“…In the field, researchers usually observe intertidal animals at low tide and only get a "snapshot" of predation at that time (Fairweather & Underwood 1983). Yet predator foraging behavior may depend on tidal conditions (Menge 1974, Checon et al 2020. Prey-handling times also differ among prey species (Fairweather & Underwood 1983, Carroll & Wethey 1990, Creswell & Mclay 1990, Soto et al 2004, Yamamoto 2004, López et al 2010, Meirelles & Matthews-Cascon 2016, Song et al 2019, because prey with more defensive morphologies require longer handling times, which in turn can bias field observations (Fairweather & Underwood 1983).…”