“…Highly abrasive diet typical for grazers causes low and blunt wear relief on the teeth. This method has been successfully applied to fossil ungulate teeth (e.g., Kaiser, 2004;Mihl-4 bachler and Solounias, 2006;Rivals et al, 2007;Croft and Weinstein, 2008;DeMiguel et al, 2008;Kahlke and Kaiser, 2011;Mihlbachler et al, 2011) and is grounded in studies of modern ungulates of known diet (e.g., Fortelius and Solounias, 2000;Clauss et al, 2007). Morphological adaptations improving wear resistance of herbivorous mammal teeth, such as increased molar crown height (hypsodonty), reflect the conditions under which they evolved and have been shown to be adaptive both to environmental factors (such as precipitation and erosion rates, which regulate the accumulation of mineral particles on plant material), as well as to dietary factors (abrasive food items, mainly grass).…”