“…An estimated 40% or more of the Atlantic Ocean floor was formed by interlinked processes of asymmetric extension, magmatism, and detachment faulting resulting in exposure of mantle and lower crustal rocks in OCCs (Dick et al, 2008;Cannat et al, 2006;Smith et al, 2006Smith et al, , 2008Ildefonse et al, 2007;Escartín et al, 2003;Tucholke et al, 2008). OCCs incorporate highly reactive olivine-rich rocks that interact with seawater over a range of temperatures to produce both high-and low-temperature hydrothermal systems (Douville et al, 1997(Douville et al, , 2002Charlou et al, 1998Charlou et al, , 2002Sagalevich et al, 2000;Kelley et al, 2005;McCaig et al, 2007). Chemical exchange between the mantle and the hydrosphere supports abundant microbial communities in regions of focused fluid discharge in both types of systems (Schrenk et al, 2004;Takai et al, 2004;Nercessian et al, 2005;Brazelton et al, 2006;Campbell et al, 2006;Perner et al, 2007).…”