“…The alternative hypothesis, involving island-hopping across the Indian Ocean, should also be considered, as recent investigations have highlighted the critical role archipelagos that are currently immersed but were likely above water when the sea-level was low (but see Sheth et al, 2003 andThébaud et al, 2009) in the biogeography of the southwestern Indian Ocean region (Cuenca et al, 2008;Le Péchon et al, 2015;Renner et al, 2010;Strijk et al, 2014). An Asian origin of the Southwestern Indian Ocean Polystichum species is in line with the view that a large part of biota of Madagascar and nearby islands share strong biogeographical affinities with Asia (Warren et al, 2010;Yoder and Nowak, 2006).…”