“…Ichnologic assessment is important in aquifer and reservoir characterization,however, most studies have been conducted in siliciclastic units (e.g., Keswani and Pemberton, 2007;Tonkin et al, 2010;Gingras et al, 2012). Bioturbation commonly decreases porosity and permeability in both siliciclastic and carbonate aquifers that contain significant intergranular flow (e.g., Gingras et al, 2004;.Karstic aquifers such as the Edwards and Trinity Groups; however, cannot transmit significant quantities of fluid through interparticle porosityand rely on bioturbation to create porosity and lateral fluid pathways (e.g., Mathews, 1967;Gingras et al, 1999;Cunningham and Sukop, 2011;Baniak et al 2013;Golab et al, 2015).Although previous authors have conducted ichnological assessment of some carbonate hydrocarbon reservoirs (e.g., Gingras et al, 1999Gingras et al, , 2004La Croix et al, 2011;Baniak et al, 2013), thebiogenic aspect of karstic aquifers has been overlooked in the literature (Cunningham et al, 2009;Sukop, 2011, 2012;Golab et al, 2015;Clark et al, 2016). The GRL is a dual-permeability system and absent fracture and karstic porosity, ichnofossils with coarse-grained infill are the primary fluid pathways throughout most strata and are a significant factor in directing water between major faults and karstic features.…”