“…The roughness of a surface in contact with a moving fluid can have substantial effects on the character of flow near that surface and on the nature of the shear stress in the boundary layer, which determines one component of drag (Schetz, 1993; Smits, 2000; Vogel, 1994). Effects of surface roughness are extremely complex, and depend on the scale of the roughness, orientation of surface elements, fluid velocity, Reynolds number, and the movement of the surface (Raayai‐Ardakani & McKinley, 2019). This latter feature, how any particular surface moves in flow, is particularly difficult to correlate with specific effects on the boundary layer: analyses of moving biological surfaces and how motion affects near‐surface flow dynamics have proven to be challenging to perform (Anderson, McGillis, & Grosenbaugh, 2001; Eloy, 2013; Lauder et al, 2016; Taneda & Tomonari, 1974; Yanase & Saarenrinne, 2015).…”