A large fraction of the world grasslands and savannas are undergoing a rapid shift from herbaceous to woody-plant dominance. This land-cover change is expected to lead to a loss in livestock production (LP), but the impacts of woody-plant encroachment on this crucial ecosystem service have not been assessed. We evaluate how tree cover (TC) has affected LP at large spatial scales in rangelands of contrasting social-economic characteristics in the United States and Argentina. Our models indicate that in areas of high productivity, a 1% increase in TC results in a reduction in LP ranging from 0.6 to 1.6 reproductive cows (Rc) per km 2 . Mean LP in the United States is 27 Rc per km 2 , so a 1% increase in TC results in a 2.5% decrease in mean LP. This effect is large considering that woody-plant cover has been described as increasing at 0.5% to 2% per y. On the contrary, in areas of low productivity, increased TC had a positive effect on LP. Our results also show that ecological factors account for a larger fraction of LP variability in Argentinean than in US rangelands. Differences in the relative importance of ecological versus nonecological drivers of LP in Argentina and the United States suggest that the valuation of ecosystem services between these two rangelands might be different. Current management strategies in Argentina are likely designed to maximize LP for various reasons we are unable to explore in this effort, whereas land managers in the United States may be optimizing multiple ecosystem services, including conservation or recreation, alongside LP.global change | food production | social-ecological systems G rasslands, shrublands, and savannas, collectively termed "rangelands," constitute about 50% of the Earth's land surface (1). Although characterized by low yet highly variable annual rainfall, these areas provide 30-35% of terrestrial net primary productivity (NPP) (2), contain >30% of the world's human population, and support the majority of the world's livestock production (LP) (3, 4). Besides LP, rangelands also provide a variety of other ecosystem services, including fiber production, carbon sequestration, maintenance of the genetic library (conservation), and recreation (5).One of the most striking land-cover changes in rangelands worldwide over the past 150 y has been the proliferation of trees and shrubs at the expense of perennial grasses (6). In the United States, nonforest lands undergoing woody-plant encroachment are now estimated to cover up to 335 million ha (40% of the coterminous United States) (7) and the increase in woody cover ranges from 0.5% to 2% per y (8). The causes of this vegetation change are debated and the main potential drivers include intensification of livestock grazing, changes in climate and fire regimes, the introduction of nonnative woody species, and declines (natural and human induced) in the abundance of browsing animals (9-12). Historical increases in atmospheric nitrogen deposition and atmospheric carbon dioxide concentration have also been suggested to play ...