Fossil assemblages of the Ordovician to Devonian successions of Japan suggest complex temporal, environmental and geographical controls on their biogeographical signature. Thus, limited similarity at the species-level between the trilobite, brachiopod and ostracod faunas of the South Kitakami, Hida-Gaien and Kurosegawa terranes in part reflects the sporadic stratigraphic distribution of shelly fauna within these terranes. As a result, and with the exception of corals and pan-tropical radiolarians, species-level similarities are greater with other regions of East