“…In subalpine grassland in the eastern Pyrenees (Andorra), heavy grazing increased species richness and diversity, but decreased forage quality and production (Komac et al, 2014). Overgrazing has profound effects on important ecosystem characteristics, such as water erosion (Cerdà and Lavee, 1999), soil water content (SWC) (Lin et al, 2010), soil organic carbon (SOC) (Su et al, 2006;Costa et al, 2015), total nitrogen (TN) (Augus- tine and Frank, 2001;Lin et al, 2010;Hirobe et al, 2013), plant species richness and diversity (Gibson, 1988;Hickman et al, 2004;Ren et al, 2012;Angassa, 2014), belowground bud bank (Qian et al, 2014) and ecosystem stability (Su et al, 2006;Ren et al, 2012). However, the effects of livestock grazing on soil heterogeneity have not been consistent -with some studies showing that overgrazing increased soil heterogeneity of TN, SOC and fine soil proportion (Schlesinger et al, 1990;Su et al, 2006), while other studies reported that soil heterogeneity and vegetation diversity decreased from a patchy to a homogeneous distribution with increased grazing intensity (e.g., Zhao et al, 2011).…”