“…Moreover, the relative abundances of the ααα-20R C 27 , C 28 and C 29 regular steranes (63%, 22% and 15% respectively) indicate a predominantly marine input, corroborated by the additional presence of the more unusual C 30 steranes, which are indicative of a marine source 47 . Interestingly, secosteranes, specifically the C 27 , C 28 and C 29 8,14-secosteranes, were identified; these are believed to derive from sponge lipid membranes and their symbiotic bacteria, as previously reported in marine-derived fossil fuels 49 . The decreasing C 31 -C 35 homohopanes and the diasteranes, being in similar abundance to the regular steranes, also suggests a clastic input for the lignite 54 .…”