“…Earlier research indicated that the chromatin of crustacean sperm is not highly packaged due to a lack of basic proteins. However, several studies on Cancer pagurus (Kurtz, Martínez-Soler, Ausió, & Chiva, 2008), Cancer magister (Kurtz et al, 2008), Maja brachydactyla (Kurtz, Ausió, & Chiva, 2009), Portunus pelagicus (Stewart et al, 2010), Fenneropenaeus chinensis (Ge et al, 2011), Astacus (Niksirat, James, Andersson, Kouba, & Kozák, 2015), and Eriocheir sinensis (Li et al, 2017;Wu et al, 2016;Wu, Kang, Guo, Mu, & Zhang, 2015;Zhang et al, 2016) have indicated that these decapod spermatozoal nuclei have a common characteristic, namely, that they contain parts of histones and some histone modifications. Because of the diversity of crustacean species, an integrated understanding of the origin of this special decapod sperm nuclei has not been achieved.…”