“…For example, lower FIDs in sheltered versus exposed habitats suggest that prey perceive less risk when shelter is available (de Boer van K E Y W O R D S antipredator behaviour, economic escape theory, hatchery, migration, movement, predation risk, speed Breukelen, Hootsmans, & van Wieren, 2004). Likewise, a longer time to emerge from refuge when more conspecifics are present suggests that prey perceive that situation to be riskier, possibly due to predator aggregation in response to prey density (Pezner, Lim, Kang, Armenta, & Blumstein, 2017). The general question escape theory models ask is as follows: how much predation risk do prey perceive?…”