Regional climate responses to large-scale forcings, such as precessional changes in solar irradiation and increases in anthropogenic greenhouse gases, may be nonlinear as a result of complex interactions among earth system components. Such nonlinear behaviors constitute a major source of climate "surprises" with important socioeconomic and ecological implications. Paleorecords are key for elucidating patterns and mechanisms of nonlinear responses to radiative forcing, but their utility has been greatly limited by the paucity of quantitative temperature reconstructions. Here we present Holocene July temperature reconstructions on the basis of midge analysis of sediment cores from three Alaskan lakes. Results show that summer temperatures during 10,000-5,500 calibrated years (cal) B.P. were generally lower than modern and that peak summer temperatures around 5,000 were followed by a decreasing trend toward the present. These patterns stand in stark contrast with the trend of precessional insolation, which decreased by ∼10% from 10,000 y ago to the present. Cool summers before 5,500 cal B.P. coincided with extensive summer ice cover in the western Arctic Ocean, persistence of a positive phase of the Arctic Oscillation, predominantly La Niña-like conditions, and variation in the position of the Alaskan treeline. These results illustrate nonlinear responses of summer temperatures to Holocene insolation radiative forcing in the Alaskan sub-Arctic, possibly because of state changes in the Arctic Oscillation and El Niño-Southern Oscillation and associated land-atmosphere-ocean feedbacks.Chironomidae | paleotemperature | climate change | high latitudes | land-ocean-atmosphere interaction A lthough radiative forcing by greenhouse gases is spatially uniform, temperature trajectories have displayed high variability both spatially and temporally during the 20th and 21st centuries. Understanding the patterns and causes of this variability is paramount to anticipating the impacts of climate change at local to regional scales. Among the major contributing factors is nonlinearity in regional temperature responses to large-scale climate forcings. Such nonlinearity may result from land-ocean feedbacks to atmospheric warming and atmosphere-ocean teleconnections associated with climate modes such as the Arctic Oscillation/North Atlantic Oscillation (AO/NAO) (1). For example, rising global mean temperatures are projected to increase prevalence of the AO/NAO positive phase by the end of this century. This change could accentuate the contrast of warming over Eurasia and interior Canada with cooling over northeastern Canada.Paleorecords can help elucidate mechanisms leading to nonlinear and spatially heterogeneous responses of temperature to radiative forcing. During the Holocene, high-latitude insolation varied markedly with the precessional cycle; for example, July short-wave radiation declined by ∼50 W m −2 at 65°N latitude throughout the Holocene (2). Paleoclimate data and model simulations have demonstrated the dominant effect of d...