We thank Clarke (2019) for his defense of Australopithecus prometheus. Clarke disagrees with our assessment (Hawks and Berger, 2019) of what Dart purported. In his view, Dart's statement about the more extensive dietary and sutural system attributed to MLD 1 are sufficient to make the species name Au. prometheus available under the ICZN. Clarke further argues that we are incorrect to read Dart's description as conditional. In his view, Au. prometheus is not a nomen nudum.Our commentary did refer to the possibility that others might find that Dart's definition of Au. prometheus was not deficient in the ways we have suggested. In this event, Au. prometheus is a nomen dubium. We regard this to be an uncontroversial statement. After Robinson (1954) nearly every specialist has regarded Plesianthropus transvaalensis as a junior synonym of Au. africanus. MLD 1 is insufficient to support the diagnosis of Au. prometheus as distinct from either.In his letter, Clarke refers to two features that in his opinion substantiate a difference between MLD 1 and Au. africanus, for which he takes Sts 5 as an example. These are: "the large brain size of MLD 1 and the vertical sides of its vault with parietal bossing." Here we examine both features in turn.Dart viewed MLD 1 to be of the same endocranial volume he would expect for an adult version of the Taung specimen of Au.africanus. In his assessment (Dart, 1948:273), "the endocranial cast of this adult occiput confirms and corroborates the evidence of cerebral expansion and intellectual superiority furnished over 20 years ago by the Taungs endocranial cast." Holloway (1972)