“…Proline is an important osmotic regulator when plants respond to environment change. Usually plants are influenced to external stresses (Peng et al 2019(Peng et al , 2020, all kinds of osmoregulation substances, such as proline, betaine, and monosaccharides which will accumulate to deal with the stress through osmotic adjustment (Serrano 1996). Proline accumulation is influenced by drought, salinity, metals, and biotic stresses (Kavi et al 2005, Wang et al 2007.…”