“…Rice breeding programs need to incorporate NuUE traits as this helps resource-poor farmers to save on fertilizer, and allows maximization of valuable resources to increase profitability. To date, several morphological and agronomic traits have been identified, and they can be used as indicators of nitrogen (N) [ 52 , 64 , 65 , 66 ], phosphorus (P) [ 7 , 38 , 67 , 68 , 69 , 70 ], and potassium (K) [ 16 , 39 , 40 , 71 , 72 , 73 ] deficiency tolerance under both field and hydroponic conditions. Using different genetic backgrounds of mapping populations, such as RILs, NILs, BILs, ILs, CSSLs, DHs, and BC 2 F 4 , there are several reports on QTL identification with independent studies of low-N, -P, and -K conditions.…”