“…Recently, artificial neural networks (ANNs), inspired by the brain, have been actively researched and developed [ 1 ]. Particular attention has been paid to spiking neural net-works (SNNs) that utilize biological spike timing–dependent plasticity (STDP) rules to function as event-driven systems for unsupervised learning [ 2 , 3 , 4 , 5 , 6 , 7 , 8 , 9 , 10 ]. Since such SNNs pursue a more bio-plausible direction than other ANNs do, it is necessary to explore the signal transmission mechanisms of biological neurons and synapses to gain a deeper understanding of these SNNs.…”