When associations between certain visual stimuli and particular actions are learnt, those stimuli become capable of automatically and unconsciously activating their associated action plans. Such sensorimotor priming is assumed to be fundamental for efficient responses, and can be reliably measured in masked prime studies even when the primes are not consciously perceived. However, when the delay between prime and target is increased, reversed priming effects are often found instead (the negative compatibility effect, NCE). The main accounts of the NCE assume that it too is a sensorimotor phenomenon, predicting that it should occur only when the initial positive priming phase also occurs. Alternatively, reversed priming may reflect a perceptual process entirely independent from positive motor priming (which is simply evident at a different temporal delay), in which case no dependency is expected between the NCE and positive priming. We tested these predictions while new sensorimotor associations were learnt, and when learnt associations were suddenly reversed. We found a remarkable symmetry between positive and reversed priming during all such learning phases, supporting the idea that reversed priming represents a sensorimotor process that is contingent on, and automatically follows, the positive priming phase. We discuss also whether the NCE mechanism is subject to a trigger threshold.