“…1) could not have occurred without an increase in source activity per unit area, which has both a rate (i.e., canopy photosynthesis, g CO 2 m –2 GA d –1 , where GA is ground area) and a time component (principally the duration of kernel filling; Egli, 2011). The importance of kernel number in the increase in yield (Carlone and Russell, 1987; Russell, 1991; Tollenaar et al, 1992; Echarte et al, 2000; Li et al, 2011) suggests that the rate component was more important than the time component (longer kernel filling period). Traits listed by Duvick et al (2004a) and Duvick (2005) that contributed to increased productivity included smaller tassels, erect leaves, increased stay‐green characteristics of the leaves during kernel filling, less lodging, and improved disease and insect resistance.…”