“…The long-term sustained coexistence of yellow pine species and the SPB suggests that fire and SPB mortality historically interacted to perpetuate yellow pine forests. In contemporary forests, dead trees resulting from SPB infestations create heavy fuel loads that may result in more intense fires that kill more trees and reduce stand density (Kuykendall, 1978;Knebel and Wentworth, 2007). However, Schowalter et al (1981) suggested that upland pine forests prior to fire suppression were not susceptible to heavy SPB outbreaks because low stand density resulted in reduced tree stress, likely due to reduced competition (Coulson, 1979) and reduced effectiveness of pheromone communication by the SPB (Fares et al, 1980;Thistle et al, 2004).…”