The proto-Paratethys was a vast east-west oriented inland sea within mid-latitudes during the Cretaceous to Paleogene times, which extended across Eurasia from western Europe to the Tarim Basin in western China (Kaya et al., 2018;Radionova et al., 2003; Figure 1a). From the early Eocene to the Oligocene, the sea experienced longterm regression, superimposed by major transgression-regression cycles due to tectonic and/or eustatic effects (Kaya et al., 2018;Meijer et al., 2019;Radionova et al., 2003). At its eastern end, the transgressive and regressive intervals of the proto-Paratethys are clearly recorded in the Tarim Basin and coincide with the humid and arid phases, respectively in the Qaidam and Xining basins, demonstrating the role of the proto-Paratethys as an important moisture source for the Asian interior (Bosboom et al., 2014;Kaya et al., 2018). Modeling studies also suggest that the westward retreat of the proto-Paratethys could strengthen the Asian monsoons by enlarging the land-sea thermal contrast (Ramstein et al., 1997;Z. Zhang et al., 2007).Besides, the sea level fluctuations of the proto-Paratethys could alter the seaways between the proto-Paratethys and the adjacent oceans (Iakovleva, 2011;Kaya et al., 2018;Radionova et al., 2003). During the early Paleogene, there are two potential connections of the proto-Paratethys with the mid-and high-latitude oceans: one is westward through the Polish Lowland Basin to the North Sea and the North Atlantic; the other is northward through the Turgai Strait to the West Siberia Sea and the Arctic (Figure 1a). The correlation of sediment facies across the proto-Paratethys area suggests that in the late Paleocene and the early Eocene, the two open connections ensured an extensive water exchange between the proto-Paratethys and the Arctic (Iakovleva, 2011;Radionova et al., 2003). Further studies revealed largely similar microfossil assemblages between the proto-Paratethys and the North Sea, confirming that these two basins were indeed connected during the early Eocene (Deprez et al., 2015;King et al., 2013). It seems that the exchange of water masses between the proto-Paratethys and the Arctic via the