“…As a world crop, cotton has been cultivated for at least 3000 yr (Lee & Fang, 2015). Tetraploid upland cotton ( Gossypium hirsutum L.) is the major species grown in the United States (Bellaloui & Turley, 2013; Tewolde et al., 2018) and worldwide (Amin et al., 2018; Basal et al., 2019; Kuang et al., 2018; Zhang, Zhang, Pu, Zhang, & Ou, 2019). Another tetraploid species known as pima cotton ( G. barbadense L.) is also cultivated (Cornish, Radin, Turcotte, Lu, & Zeiger, 1991) with the characteristics of longer, finer, and stronger fibers (Jenkins, McCarty, Gutierrez, Hayes, & Jones, 2013).…”