“…On the other hand Butler (1967) found that the intensity of an odour trail and its attraction to potential foragers vary with the amount and quality of the food supply. It is quite clear that in several ant species, scouts initially use visual orientation to randomly search for food independently; the rewarded scout then communicates the discovered food by laying pheromonal trail from the source to the nest (Carthy, 1951;Wilson, 1965;Leuthold, 1968;SzlepFessel, 1970;Holldobler, 1971Holldobler, , 1976Moglich and Holldobler, 1975;Jaffe and Howse, 1979).…”