Climate trends on timescales of 10s to 100s of millions of years are controlled by changes in solar luminosity, continent distribution, and atmosphere composition. Plate tectonics affect geography, but also atmosphere composition through volcanic degassing of CO 2 at subduction zones and midocean ridges. So far, such degassing estimates were based on reconstructions of ocean floor production for the last 150 My and indirectly, through sea level inversion before 150 My. Here we quantitatively estimate CO 2 degassing by reconstructing lithosphere subduction evolution, using recent advances in combining global plate reconstructions and present-day structure of the mantle. First, we estimate that since the Triassic (250-200 My) until the present, the total paleosubduction-zone length reached up to ∼200% of the present-day value. Comparing our subduction-zone lengths with previously reconstructed ocean-crust production rates over the past 140 My suggests average global subduction rates have been constant, ∼6 cm/y: Higher ocean-crust production is associated with longer total subduction length. We compute a strontium isotope record based on subduction-zone length, which agrees well with geological records supporting the validity of our approach: The total subduction-zone length is proportional to the summed arc and ridge volcanic CO 2 production and thereby to global volcanic degassing at plate boundaries. We therefore use our degassing curve as input for the GEOCARBSULF model to estimate atmospheric CO 2 levels since the Triassic. Our calculated CO 2 levels for the mid Mesozoic differ from previous modeling results and are more consistent with available proxy data.paleoclimate | carbon cycle | geodynamic V olcanism forms the most efficient mechanism to transfer carbon (C) from the mantle to the ocean-atmosphere system, the exogenic reservoir. At midocean ridges, mantle upwelling occurs between two diverging plates. At subduction zones, sinking oceanic plates lead to arc volcanism (1). A third type of volcanism, driven by mantle plumes, leads occasionally to large igneous provinces (LIPs) that typically last a few My (2, 3). Longterm (>>5 My) changes in volcanic output thus dominantly relate to the long-term plate tectonic processes. Constraints on volcanism-related degassing of CO 2 at subduction zones and midocean ridges hence derive from studies on lithosphere production/consumption (3-6). The quality of these constraints determines the mechanistic underpinning of climatic changes on timescales of 10 7 to 10 8 y. This is because on such timescales, volcanism is the major source of CO 2 to the atmosphere and thereby drives climate and the critical feedbacks in the carbon cycle, such as weathering. To date, estimates of CO 2 degassing through plate tectonic processes for the last 150 My are based on reconstructions of ocean-floor production (7), and for older periods indirect inferences were made through sea level inversion (8). An increasing body of work reveals major uncertainties in eustatic sea level reconstructi...