“…Hole-nesting birds are considered as one of the most susceptible groups to such impacts due to their strong dependence on trees (Angelstam & Mikusiński 1994, Martin & Eadie 1999, Imbeau et al 1999. It has been suggested that the availability of nesting holes limits the population density of such birds in many managed forests (von Haartman 1957, Johnson 1994, Newton 1994, Semel & Sherman 2001, Twedt & Henne-Kerr 2001, Pöysa & Pöysa 2002. Since the 1990s, with the emergence of ecological forestry, there is an increasing need to improve our understanding of biodiversity and its maintenance in natural forests, so that such knowledge can be incorporated into management guidelines (Hansen et al 1991, Fujimori 2001.…”