“…1 2 QIAN LIU, SHUANG LIU AND KING-YEUNG LAM ASYMPTOTIC SPREADING OF INTERACTING SPECIES 3 Nevertheless, the exact formula of c 2 remained open.In the bistable case a, b > 1 with appropriate initial conditions, the spreading problem was studied by Carrère [9], who showed that solutions of (1) exhibit two moving interfaces connecting, starting from the right, (0, 0) to (0, 1) to (1, 0). The first interface moves with the expected speed of c 1 = 2 √ dr, whereas the second interface moves with speed c 2 which is the speed of the unique traveling wave solution connecting (0, 1) to (1, 0).The monostable case 0 < a < 1 < b, which is closely related to our problem, was considered by Girardin and the last author [18]. By a delicate construction of piecewise smooth super-and sub-solutions for (1), it was shown that while the faster species spreads at the expected speed c 1 = 2 √ dr, the spreading speed c 2 of the slower species depends on c 1 in a non-trivial way, and is a nonlocally determined quantity in general (see Subsection 1.1 for details).…”