Eight years' fluctuations of fruit production were studied in a warn temperate broadleaved forest on Yakushima Island, southern Japan. Eight tree species characterized by sapfruits or capsules were studied, and all of them were regarded as animal dispersal Kelly, 1988) and predator avoidance hypothesis (Jarlzen,1971;Silvertown, 1980) are applicable for many species.The iesource matching hypothesis and the predator avoidance hypothesis belong to the different categories of explanation: the former is based on a proximate factor and the latter on an ultimate factor.Resource matching hypothesis mentions that climate is a primary factor: the good photosynthetic condition of a tree leads to mast fruiting and unusual weather restricts the level of investments to fruits, so that the fluctuation of rich or poor harvest becomes uniform within many individuals. Biennial fruiting is well known in the fruit horticulture (Hoblyn et al., !936), and the fluctuation of fruit production in orchards synchronizes entirely regional, or more widely, often in all Japan (Ito er al ', 1956)' For an aim to reduce annual harvest fluctuation, physiological mechanism offruit trees has been studied well (Nakagawa, t982). In cases of various fruit trees, the accumulation of starch is related to the decision of flower bud quantity (Davis, 1931; ogaki et al., !963; Sobajima, 1967).Predator avoidance hypothesis explains that the steep fluctuation of fruiting is an adaptation as many plants avoid seed predators in mast year by controlling predator's number low in poor harvest