“…Previous works on Permian Karoo black shales provide sedimentological and palaeontological evidence for their marine origin (e.g., Haughton et al, 1953;du Toit, 1954;Rilett, 1963;Hart, 1964Hart, , 1969Hart, , 1970Ryan, 1968;McLachlan and Anderson, 1973;Teichert and Rilett, 1974;Stanistreet et al, 1980;Oelofsen, 1987;Visser, 1987Visser, , 1989Visser, , 1992Visser, , 1993Visser, , 1994Visser and Young, 1990;Smith et al, 1993;Johnson et al, 1997Johnson et al, , 2006Scheffler et al, 2006;Buatois et al, 2010;Götz, 2015). Despite this, there is ongoing controversy and discussion concerning the marine or lacustrine origin in the context of the postglacial development of the Karoo Basin (e.g., Cole and McLachlan, 1991;Faure and Cole, 1999;Herbert & Compton, 2007;Chukwuma and Bordy, 2016;Schulz et al, 2016). This discussion includes the palaeoecology of the famous Permian mesosaurs (Oelofsen, 1981;Oelofsen and Araújo, 1983), interpretations ranging from freshwater, brackish, coastal, lagoonal, hypersaline to normal marine (see Modesto, 2006;Piñeiro et al, 2012;Silva et al, 2017 and references therein).…”