Let n ě 2 and r P t1, ¨¨¨, n ´1u be integers, M be a compact smooth Kähler manifold of complex dimension n, E be a holomorphic vector bundle with complex rank r and equipped with an hermitian metric h E , and L be an ample holomorphic line bundle over M equipped with a metric h with positive curvature form. For any d P N large enough, we equip the space of holomorphic sections H 0 pM, E b L d q with the natural Gaussian measure associated to h E , h and its curvature form. Let U Ă M be an open subset with smooth boundary. We prove that the average of the pn ´rq-th Betti number of the vanishing locus in U of a random section s of H 0 pM, E b L d q is asymptotic to `n´1 r´1 ˘dn ş U c 1 pLq n for large d. On the other hand, the average of the other Betti numbers are opd n q. The first asymptotic recovers the classical deterministic global algebraic computation. Moreover, such a discrepancy in the order of growth of these averages is new and constrasts with all known other smooth Gaussian models, in particular the real algebraic one. We prove a similar result for the affine complex Bargmann-Fock model.