“…Like many other ancient lakes, the Titicaca has a complex palaeo-environmental history (Cristescu et al, 2010). Thus, the different Altiplano basins conformed a single hydrological unit from the Early to Middle Pleistocene (Baker & Fritz, 2015;Nunnery et al, 2019), when water level experienced at least three major expansions (high stands) of up to 114, 104 and 54 m above present lake level (Fornari et al, 2001). Furthermore, the lake has passed through several periods of retraction coincident with global interglacial periods, when its level FIGURE 1 A, map of South America showing location of the Andean Altiplano; B, approximate area of the Altiplano showing placement of its main water bodies; C, sampling sites placed outside Lake Titicaca; D, sampling sites at Lake Titicaca itself.…”