Abstract. The impacts of different emission levels and climate change conditions to landscape-scale natural vegetation could have large repercussions for ecosystem services and environmental health. We forecast the risk-reduction benefits to natural landscapes of lowering business-as-usual greenhouse gas emissions by comparing the extent and spatial patterns of climate exposure to dominant vegetation under current emissions trajectories (Representative Concentration Pathway, RCP8.5) and envisioned Paris Accord target emissions (RCP4.5). This comparison allows us to assess the ecosystem value of reaching targets to keep global temperature warming under 2°C. Using 350,719 km 2 of natural lands in California, USA, and the mapped extents of 30 vegetation types, we identify each type's current bioclimatic envelope by the frequency with which it occupies current climate conditions. We then map the trajectory of each pixel's climate under the four climate futures to quantify areas expected to fall within, become marginal to (outside a 95% probability contour), or move beyond their current climate conditions by the end of the 21st century. In California, these four future climates represent temperature increases of 1.9-4.5°C and a À24.8 to +22.9% change in annual precipitation by 2100. From 158,481 to 196,493 km 2 (45-56%) of California's natural vegetation is predicted to become highly climatically stressed under current emission levels (RCP8.5) under the drier and wetter global climate models, respectively. Vegetation in three California ecoregions critical to human welfare, southwestern CA, the Great Valley, and the Sierra Nevada Mountains, becomes >50% impacted, including 68% of the lands around Los Angeles and San Diego. However, reducing emissions to RCP4.5 levels reduces statewide climate exposure risk by 86,382-99,726 km 2 . These projections are conservative baseline estimates because they do not account for amplified drought-related mortality, fires, and beetle outbreaks that have been observed during the current five-year drought. However, these results point to the landscape benefits of emission reductions.